We may consequently state the fundamental theorem of Natural Selection in the form : The rate of increase in fitness of any organism at any time is equal to its genetic variance in fitness at that time. (Ronald Fisher)

We may consequently state the fundamental theorem of Natural Selection in the form : The rate of increase in fitness of any organism at any time is equal to its genetic variance in fitness at that time.

Ronald Fisher

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equal fitness form fundamental increase natural organism rate selection state theorem time variance

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